I'm not sure if it works since I have never changed this and there are a few "unconnected" UI elements in Fusion that do nothing. But the general idea is, that Fusion saves the result of several nodes in the RAM. This is the green line in the playbar. You can disable it completely (which I wouldn't advise when you're working on a composition) or set a threshold that it only saves the results to RAM if the playback speed is lower than the specified value.
If I would ever use this I would usually set this to the project frame rate (or just slightly lower).
But in general I wouldn't touch this at all if your not running out of RAM constantly.